Region: CIP
Geographic Zones: Eastern Indonesia (incl. East Timor) - Zone 4 » Sulawesi - Zone 43 » North Sulawesi - Zone 431 » Bunaken National Marine Park and Manado - Zone 4313 » Tiwoho - Zone 43130067
Survey Types: Species & Abundance, Species Only
Dates: 1/1/93 - 12/9/24
Total Surveys: 2
#Species Reported: 209
Average Species Reported on a Survey by Expert Surveyors: 147.00
Average Species Reported on a Survey by Novice Surveyors: 0
Survey Type: SA = Species & Abundance; SO = Species Only - How to interpret REEF data?
Click a column header to sort by that column. Hold shift and click to sort by multiple columns
Surveys | Bottom Time | |||||
---|---|---|---|---|---|---|
Expert | Novice | |||||
Code | Name | SA | SO | SA | SO | (H:M) |
4 | Eastern Indonesia (incl. East Timor) | 2 | 0 | 0 | 0 | 2:24 |
43 | Sulawesi | 2 | 0 | 0 | 0 | 2:24 |
431 | North Sulawesi | 2 | 0 | 0 | 0 | 2:24 |
4313 | Bunaken National Marine Park and Manado | 2 | 0 | 0 | 0 | 2:24 |
43130067 | Tiwoho | 2 | 0 | 0 | 0 | 2:24 |
TOTALS | 2 | 0 | 0 | 0 | 2:24 |
Species
%SF = Sighting Frequency; DEN = Density Score - How to interpret REEF data?
Bar length corresponds to sighting frequency
Color saturation corresponds to density score
Click + to display species image and additional information.
Bar length corresponds to sighting frequency
Color saturation corresponds to density score
Click + to display species image and additional information.
Ambon Damsel - Damselfish
(Pomacentrus amboinensis)
(Pomacentrus amboinensis)
Bicolor Chromis - Damselfish
(Pycnochromis margaritifer)
(Pycnochromis margaritifer)
Bicolor Cleaner Wrasse - Wrasse
(Labroides bicolor)
(Labroides bicolor)
Black-backed Butterflyfish - Butterflyfish
(Chaetodon melannotus)
(Chaetodon melannotus)
Black-saddled Toby - Puffer
(Canthigaster valentini)
(Canthigaster valentini)
Blackbar Chromis - Damselfish
(Pycnochromis retrofasciatus)
(Pycnochromis retrofasciatus)
Blackbelt Hogfish - Wrasse
(Bodianus mesothorax)
(Bodianus mesothorax)
Blackeye Thicklip - Wrasse
(Hemigymnus melapterus)
(Hemigymnus melapterus)
Blacklip Butterflyfish - Butterflyfish
(Chaetodon kleinii)
(Chaetodon kleinii)
Blackstreak Surgeonfish - Surgeonfish
(Acanthurus nigricauda)
(Acanthurus nigricauda)
Blackstripe Cardinalfish - Cardinalfish
(Ostorhinchus nigrofasciatus)
(Ostorhinchus nigrofasciatus)
Blacktail Snapper - Snapper
(Lutjanus fulvus)
(Lutjanus fulvus)
Bleeker's Parrotfish - Parrotfish
(Chlorurus bleekeri)
(Chlorurus bleekeri)
Bluesided Fairy Wrasse - Wrasse
(Cirrhilabrus cyanopleura)
(Cirrhilabrus cyanopleura)
Bluestreak Cleaner Wrasse - Wrasse
(Labroides dimidiatus)
(Labroides dimidiatus)
Bluntheaded Wrasse - Wrasse
(Thalassoma amblycephalum)
(Thalassoma amblycephalum)
Bridled Monocle Bream - Coral (Monocle) Bream
(Scolopsis bilineata)
(Scolopsis bilineata)
Brushtail Tang - Surgeonfish
(Zebrasoma scopas)
(Zebrasoma scopas)
Charcoal Damsel - Damselfish
(Pomacentrus brachialis)
(Pomacentrus brachialis)
Cheek-lined (pka Linedcheeked) Wrasse - Wrasse
(Oxycheilinus digramma)
(Oxycheilinus digramma)
Clark's Anemonefish - Damselfish
(Amphiprion clarkii)
(Amphiprion clarkii)
Crescent Wrasse - Wrasse
(Thalassoma lunare)
(Thalassoma lunare)
Darkfin Chromis - Damselfish
(Pycnochromis atripes)
(Pycnochromis atripes)
Dash-dot Goatfish - Goatfish
(Parupeneus barberinus)
(Parupeneus barberinus)
Emperor Angelfish - Angelfish
(Pomacanthus imperator)
(Pomacanthus imperator)
Floral Wrasse - Wrasse
(Cheilinus chlorourus)
(Cheilinus chlorourus)
Gorgeous Shrimpgoby - Goby
(Amblyeleotris wheeleri)
(Amblyeleotris wheeleri)
Humphead Bannerfish - Butterflyfish
(Heniochus varius)
(Heniochus varius)
Humpnose Bigeye Bream - Emperor Bream
(Monotaxis grandoculis)
(Monotaxis grandoculis)
Jewel Damsel - Damselfish
(Stegastes lacrymatus)
(Stegastes lacrymatus)
Keyhole Angelfish - Angelfish
(Centropyge tibicen)
(Centropyge tibicen)
Latent Slingjaw Wrasse - Wrasse
(Epibulus brevis)
(Epibulus brevis)
Latticed Butterflyfish - Butterflyfish
(Chaetodon rafflesi)
(Chaetodon rafflesi)
Lemon Damsel - Damselfish
(Pomacentrus moluccensis)
(Pomacentrus moluccensis)
Longnose Butterflyfish - Butterflyfish
(Forcipiger flavissimus)
(Forcipiger flavissimus)
Manybar Goatfish - Goatfish
(Parupeneus multifasciatus)
(Parupeneus multifasciatus)
Mimic Surgeonfish - Surgeonfish
(Acanthurus pyroferus)
(Acanthurus pyroferus)
Moorish Idol - Moorish Idol
(Zanclus cornutus)
(Zanclus cornutus)
Orange-lined Triggerfish - Triggerfish
(Balistapus undulatus)
(Balistapus undulatus)
Ornate Butterflyfish - Butterflyfish
(Chaetodon ornatissimus)
(Chaetodon ornatissimus)
Oval-spot Butterflyfish - Butterflyfish
(Chaetodon speculum)
(Chaetodon speculum)
Pale-tail Chromis - Damselfish
(Chromis xanthura)
(Chromis xanthura)
Papuan Toby - Puffer
(Canthigaster papua)
(Canthigaster papua)
Peacock Grouper - Seabass
(Cephalopholis argus)
(Cephalopholis argus)
Pearl-scaled Angelfish - Angelfish
(Centropyge vrolikii)
(Centropyge vrolikii)
Pearlspotted Rabbitfish - Rabbitfish
(Siganus margaritiferus)
(Siganus margaritiferus)
Pennant Bannerfish - Butterflyfish
(Heniochus chrysostomus)
(Heniochus chrysostomus)
Red and Black Anemonefish - Damselfish
(Amphiprion melanopus)
(Amphiprion melanopus)
Redbreasted Wrasse - Wrasse
(Cheilinus fasciatus)
(Cheilinus fasciatus)
Redfin Butterflyfish - Butterflyfish
(Chaetodon lunulatus)
(Chaetodon lunulatus)
Reef Lizardfish - Lizardfish
(Synodus variegatus)
(Synodus variegatus)
Reid's Damsel - Damselfish
(Pomacentrus reidi)
(Pomacentrus reidi)
Reticulated Dascyllus - Damselfish
(Dascyllus reticulatus)
(Dascyllus reticulatus)
Ribbon Eel - Moray
(Rhinomuraena quaesita)
(Rhinomuraena quaesita)
Rolland's Damsel - Damselfish
(Chrysiptera rollandi)
(Chrysiptera rollandi)
Scaly Chromis - Damselfish
(Azurina lepidolepis)
(Azurina lepidolepis)
Scaly Damsel - Damselfish
(Pomacentrus lepidogenys)
(Pomacentrus lepidogenys)
Sixbar Wrasse - Wrasse
(Thalassoma hardwicke)
(Thalassoma hardwicke)
Sixstripe Wrasse - Wrasse
(Pseudocheilinus hexataenia)
(Pseudocheilinus hexataenia)
Spiny Chromis - Damselfish
(Acanthochromis polyacanthus)
(Acanthochromis polyacanthus)
Spotted Shrimpgoby - Goby
(Amblyeleotris guttata)
(Amblyeleotris guttata)
Staghorn Damsel - Damselfish
(Amblyglyphidodon curacao)
(Amblyglyphidodon curacao)
Stonycoral Ghostgoby - Goby
(Pleurosicya micheli)
(Pleurosicya micheli)
Striped Dwarfgoby - Goby
(Eviota sebreei)
(Eviota sebreei)
Striped Fangblenny - Blenny
(Meiacanthus grammistes)
(Meiacanthus grammistes)
Striped Monocle Bream - Coral (Monocle) Bream
(Scolopsis lineata)
(Scolopsis lineata)
Striped Triplefin - Triplefin
(Helcogramma striatum)
(Helcogramma striatum)
Swarthy Parrotfish - Parrotfish
(Scarus niger)
(Scarus niger)
Talbot's Damsel - Damselfish
(Chrysiptera talboti)
(Chrysiptera talboti)
Ternate Chromis - Damselfish
(Chromis ternatensis)
(Chromis ternatensis)
Three-spot Dascyllus - Damselfish
(Dascyllus trimaculatus)
(Dascyllus trimaculatus)
Three-striped Whiptail - Coral (Monocle) Bream
(Pentapodus trivittatus)
(Pentapodus trivittatus)
Tricolor Parrotfish - Parrotfish
(Scarus tricolor)
(Scarus tricolor)
Trumpetfish - Trumpetfish
(Aulostomus chinensis)
(Aulostomus chinensis)
Twospot Bristletooth - Surgeonfish
(Ctenochaetus binotatus)
(Ctenochaetus binotatus)
Twotone Wrasse - Wrasse
(Halichoeres prosopeion)
(Halichoeres prosopeion)
Vagabond Butterflyfish - Butterflyfish
(Chaetodon vagabundus)
(Chaetodon vagabundus)
Violet Soldierfish - Squirrelfish
(Myripristis violacea)
(Myripristis violacea)
Weber's Chromis - Damselfish
(Chromis weberi)
(Chromis weberi)
Whitestreak Monocle Bream - Coral (Monocle) Bream
(Scolopsis ciliata)
(Scolopsis ciliata)
Yellow-barred Parrotfish - Parrotfish
(Scarus dimidiatus)
(Scarus dimidiatus)
Yellowfin Parrotfish - Parrotfish
(Scarus flavipectoralis)
(Scarus flavipectoralis)
Yellowstripe Goatfish - Goatfish
(Mulloidichthys flavolineatus)
(Mulloidichthys flavolineatus)
Yellowtail Damsel - Damselfish
(Neoglyphidodon nigroris)
(Neoglyphidodon nigroris)
Yellowtail Fusilier - Fusilier
(Caesio cuning)
(Caesio cuning)
Ambon Chromis - Damselfish
(Pycnochromis amboinensis)
(Pycnochromis amboinensis)
Barcheek Trevally - Jack
(Craterognathus plagiotaenia)
(Craterognathus plagiotaenia)
Barred Thicklip - Wrasse
(Hemigymnus fasciatus)
(Hemigymnus fasciatus)
Bicolor Angelfish - Angelfish
(Centropyge bicolor)
(Centropyge bicolor)
Big Longnose Butterflyfish - Butterflyfish
(Forcipiger longirostris)
(Forcipiger longirostris)
Big-scale Soldierfish - Squirrelfish
(Myripristis berndti)
(Myripristis berndti)
Bignose Unicornfish - Surgeonfish
(Naso vlamingii)
(Naso vlamingii)
Black Leopard Wrasse - Wrasse
(Macropharyngodon negrosensis)
(Macropharyngodon negrosensis)
Black Snapper - Snapper
(Macolor niger)
(Macolor niger)
Black-axil Chromis - Damselfish
(Chromis atripectoralis)
(Chromis atripectoralis)
Black-finned (aka Highfin) Snake Eel - Snake Eel
(Ophichthus altipennis (prev misidentified as O. melanochir))
(Ophichthus altipennis (prev misidentified as O. melanochir))
Black-margined Damsel - Damselfish
(Pomacentrus nigromarginatus)
(Pomacentrus nigromarginatus)
Blackspot Cleaner Wrasse - Wrasse
(Labroides pectoralis)
(Labroides pectoralis)
Blackspot Shrimpgoby - Goby
(Cryptocentrus nigrocellatus)
(Cryptocentrus nigrocellatus)
Blue-axil Chromis - Damselfish
(Pycnochromis caudalis)
(Pycnochromis caudalis)
Blue-barred Parrotfish - Parrotfish
(Scarus ghobban)
(Scarus ghobban)
Bluestreak Fusilier - Fusilier
(Pterocaesio tile)
(Pterocaesio tile)
Brick Soldierfish - Squirrelfish
(Myripristis amaena)
(Myripristis amaena)
Broom Filefish - Filefish
(Amanses scopas)
(Amanses scopas)
Brown Coralblenny - Blenny
(Atrosalarias fuscus)
(Atrosalarias fuscus)
Bullethead (Pacific) Parrotfish - Parrotfish
(Chlorurus spilurus)
(Chlorurus spilurus)
Canary Wrasse - Wrasse
(Halichoeres chrysus)
(Halichoeres chrysus)
Chevroned Butterflyfish - Butterflyfish
(Chaetodon trifascialis)
(Chaetodon trifascialis)
Clearfin Lizardfish - Lizardfish
(Synodus dermatogenys)
(Synodus dermatogenys)
Cutribbon Wrasse - Wrasse
(Stethojulis interrupta)
(Stethojulis interrupta)
Decorated Sandgoby - Goby
(Istigobius decoratus)
(Istigobius decoratus)
Disappearing Wrasse - Wrasse
(Pseudocheilinus evanidus)
(Pseudocheilinus evanidus)
Double-tooth Soldierfish - Squirrelfish
(Myripristis hexagona)
(Myripristis hexagona)
Dwarf Hawkfish - Hawkfish
(Cirrhitichthys falco)
(Cirrhitichthys falco)
Epaulette Soldierfish - Squirrelfish
(Myripristis kuntee)
(Myripristis kuntee)
Fimbriated Moray - Moray
(Gymnothorax fimbriatus)
(Gymnothorax fimbriatus)
Flagtail Grouper - Seabass
(Cephalopholis urodeta)
(Cephalopholis urodeta)
Flagtail Triggerfish - Triggerfish
(Sufflamen chrysopterus (pka S chrysopterum))
(Sufflamen chrysopterus (pka S chrysopterum))
Fourline Wrasse - Wrasse
(Stethojulis trilineata)
(Stethojulis trilineata)
Freckled Goatfish - Goatfish
(Upeneus tragula)
(Upeneus tragula)
Goldbelly Damsel - Damselfish
(Pomacentrus auriventris)
(Pomacentrus auriventris)
Golden Damsel - Damselfish
(Amblyglyphidodon aureus)
(Amblyglyphidodon aureus)
Goldsaddle Goatfish - Goatfish
(Parupeneus cyclostomus)
(Parupeneus cyclostomus)
Highfin Fangblenny - Blenny
(Petroscirtes mitratus)
(Petroscirtes mitratus)
Hinged Shrimpfish - Shrimpfish
(Aeoliscus strigatus)
(Aeoliscus strigatus)
Honeycomb Grouper - Seabass
(Epinephelus merra)
(Epinephelus merra)
Honeyhead Damsel - Damselfish
(Dischistodus prosopotaenia)
(Dischistodus prosopotaenia)
Humpback Snapper - Snapper
(Lutjanus gibbus)
(Lutjanus gibbus)
Indo-Pacific Sergeant - Damselfish
(Abudefduf vaigiensis)
(Abudefduf vaigiensis)
Iridescent Cardinalfish - Cardinalfish
(Pristiapogon kallopterus)
(Pristiapogon kallopterus)
Leopard Grouper - Seabass
(Cephalopholis leopardus)
(Cephalopholis leopardus)
Leopard Wrasse / Ornate Leopard Wrasse Complex - Wrasse
(Macropharyngodon meleagris / Macropharyngodon ornatus)
(Macropharyngodon meleagris / Macropharyngodon ornatus)
Lined Bristletooth - Surgeonfish
(Ctenochaetus striatus)
(Ctenochaetus striatus)
Longfin Dottyback - Dottyback
(Manonichthys polynemus)
(Manonichthys polynemus)
Lunar Fusilier - Fusilier
(Caesio lunaris)
(Caesio lunaris)
Many-spotted Sweetlips - Sweetlips (Grunt)
(Plectorhinchus chaetodonoides)
(Plectorhinchus chaetodonoides)
Map Puffer - Puffer
(Arothron mappa)
(Arothron mappa)
Midnight Snapper - Snapper
(Macolor macularis)
(Macolor macularis)
Nagasaki Damsel - Damselfish
(Pomacentrus nagasakiensis)
(Pomacentrus nagasakiensis)
Neon Damsel - Damselfish
(Pomacentrus coelestis)
(Pomacentrus coelestis)
Neon Dwarfgoby - Goby
(Eviota atriventris)
(Eviota atriventris)
Obscure Damsel - Damselfish
(Pomacentrus adelus)
(Pomacentrus adelus)
Onespot Snapper - Snapper
(Lutjanus monostigma)
(Lutjanus monostigma)
Oriental Sweetlips - Sweetlips (Grunt)
(Plectorhinchus vittatus)
(Plectorhinchus vittatus)
Ornate Emperor - Emperor Bream
(Lethrinus ornatus)
(Lethrinus ornatus)
Pacific Double-saddle Butterflyfish - Butterflyfish
(Chaetodon ulietensis)
(Chaetodon ulietensis)
Pale Monocle Bream - Coral (Monocle) Bream
(Scolopsis affinis)
(Scolopsis affinis)
Pearly Monocle Bream - Coral (Monocle) Bream
(Scolopsis margaritifer)
(Scolopsis margaritifer)
Philippine Damsel - Damselfish
(Pomacentrus philippinus)
(Pomacentrus philippinus)
Pink Anemonefish - Damselfish
(Amphiprion perideraion)
(Amphiprion perideraion)
Pinktail Triggerfish - Triggerfish
(Melichthys vidua)
(Melichthys vidua)
Pinstriped Wrasse - Wrasse
(Halichoeres melanurus)
(Halichoeres melanurus)
Pixy Hawkfish - Hawkfish
(Cirrhitichthys oxycephalus)
(Cirrhitichthys oxycephalus)
Porcupinefish - Porcupinefish
(Diodon hystrix)
(Diodon hystrix)
Princess Damsel - Damselfish
(Pomacentrus vaiuli)
(Pomacentrus vaiuli)
Purple Anthias - Anthias
(Pseudanthias tuka)
(Pseudanthias tuka)
Quoy's Parrotfish - Parrotfish
(Scarus quoyi)
(Scarus quoyi)
Raccoon Butterflyfish - Butterflyfish
(Chaetodon lunula)
(Chaetodon lunula)
Red Filamented Flasher - Wrasse
(Paracheilinus paineorum)
(Paracheilinus paineorum)
Red Lionfish - Scorpionfish and Lionfish
(Pterois volitans)
(Pterois volitans)
Redfin Hogfish - Wrasse
(Bodianus dictynna)
(Bodianus dictynna)
Regal Angelfish - Angelfish
(Pygoplites diacanthus)
(Pygoplites diacanthus)
Richmond's Wrasse - Wrasse
(Halichoeres richmondi)
(Halichoeres richmondi)
Ringtail Wrasse - Wrasse
(Oxycheilinus unifasciatus)
(Oxycheilinus unifasciatus)
Royal Dottyback - Dottyback
(Pictichromis paccagnellae)
(Pictichromis paccagnellae)
Ruddy Fusilier - Fusilier
(Pterocaesio pisang)
(Pterocaesio pisang)
Scissortail Fusilier - Fusilier
(Caesio caerulaurea)
(Caesio caerulaurea)
Scythe Triggerfish - Triggerfish
(Sufflamen bursa)
(Sufflamen bursa)
Shorthead Fangblenny - Blenny
(Petroscirtes breviceps)
(Petroscirtes breviceps)
Shoulderbar Goby - Goby
(Gnatholepis cauerensis)
(Gnatholepis cauerensis)
Silverstreak Shrimpgoby - Goby
(Ctenogobiops crocineus)
(Ctenogobiops crocineus)
Skinspot Pygmygoby - Goby
(Trimma halonevum)
(Trimma halonevum)
Smalltooth Jobfish - Snapper
(Aphareus furca)
(Aphareus furca)
Speckled Damsel - Damselfish
(Pomacentrus bankanensis)
(Pomacentrus bankanensis)
Split-banded Cardinalfish - Cardinalfish
(Ostorhinchus compressus)
(Ostorhinchus compressus)
Sponge Ghostgoby - Goby
(Pleurosicya spongicola)
(Pleurosicya spongicola)
Spot-banded Butterflyfish - Butterflyfish
(Chaetodon punctatofasciatus)
(Chaetodon punctatofasciatus)
Spotcheek Emperor - Emperor Bream
(Lethrinus rubrioperculatus)
(Lethrinus rubrioperculatus)
Spotfin Squirrelfish - Squirrelfish
(Neoniphon sammara)
(Neoniphon sammara)
Stareye Parrotfish - Parrotfish
(Calotomus carolinus)
(Calotomus carolinus)
Stocky Anthias - Anthias
(Pseudanthias hypselosoma)
(Pseudanthias hypselosoma)
Striped Catfish - Eel-tailed Catfish
(Plotosus lineatus)
(Plotosus lineatus)
Striped Goby - Goby
(Asterropteryx striata)
(Asterropteryx striata)
Striped Sweetlips - Sweetlips (Grunt)
(Plectorhinchus lessonii)
(Plectorhinchus lessonii)
Tailspot Squirrelfish - Squirrelfish
(Sargocentron caudimaculatum)
(Sargocentron caudimaculatum)
Target Shrimpgoby - Goby
(Cryptocentrus strigilliceps)
(Cryptocentrus strigilliceps)
Threadfin Anthias - Anthias
(Pseudanthias huchtii)
(Pseudanthias huchtii)
Toothy / Fivelined Cardinalfish - Cardinalfish
(Cheilodipterus isostigmus / Cheilodipterus quinque)
(Cheilodipterus isostigmus / Cheilodipterus quinque)
Tubelip Wrasse - Wrasse
(Labrichthys unilineatus)
(Labrichthys unilineatus)
Two-spined Angelfish - Angelfish
(Centropyge bispinosa)
(Centropyge bispinosa)
Twospot Wrasse (O. bimaculatus) - Wrasse
(Oxycheilinus bimaculatus)
(Oxycheilinus bimaculatus)
Unidentified Coralgoby spp. - Goby
(Gobiodon and Paragobiodon spp.)
(Gobiodon and Paragobiodon spp.)
Violet Demoiselle - Damselfish
(Neopomacentrus violascens)
(Neopomacentrus violascens)
Wassinki Cardinalfish - Cardinalfish
(Ostorhinchus wassinki)
(Ostorhinchus wassinki)
White-spotted Puffer - Puffer
(Arothron hispidus)
(Arothron hispidus)
Whitebelly Damsel - Damselfish
(Amblyglyphidodon leucogaster)
(Amblyglyphidodon leucogaster)
Whitecheek Surgeonfish - Surgeonfish
(Acanthurus nigricans)
(Acanthurus nigricans)
Wisata Wrasse - Wrasse
(Halichoeres binotopsis)
(Halichoeres binotopsis)
Yellow Chromis - Damselfish
(Chromis analis)
(Chromis analis)
Yellow-axil Chromis - Damselfish
(Chromis xanthochira)
(Chromis xanthochira)
Yellowbarred Jawfish - Jawfish
(Opistognathus randalli)
(Opistognathus randalli)
Yellowfin Goatfish - Goatfish
(Mulloidichthys vanicolensis)
(Mulloidichthys vanicolensis)
Yellowfin Surgeonfish - Surgeonfish
(Acanthurus xanthopterus)
(Acanthurus xanthopterus)
Yellowmask Surgeonfish - Surgeonfish
(Acanthurus mata)
(Acanthurus mata)
Yellowtail Barracuda - Barracuda
(Sphyraena flavicauda)
(Sphyraena flavicauda)
Yellowtail Fangblenny - Blenny
(Meiacanthus atrodorsalis)
(Meiacanthus atrodorsalis)
Yellowtail Tubelip - Wrasse
(Diproctacanthus xanthurus)
(Diproctacanthus xanthurus)
Yellowtail Wrasse - Wrasse
(Anampses meleagrides)
(Anampses meleagrides)
Zigzag Wrasse - Wrasse
(Halichoeres scapularis)
(Halichoeres scapularis)